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Diagram illustrating the distribution of myofilaments and accessoryproteins within asarcomere.The accessory proteins aretitin, a large elastic molecule that anchors the thick (myosin) filaments to the Z line; a-actinin, which bundles thin(actin) filaments into parallel arrays and anchors them at the Z line; nebulin, an elongated inelastic protein attached to the Z linesthatwraps around thethin filaments and assists a-actinin in anchoring the thin filament to Z lines; tropomodulin, an actin-capping protein that maintains and regulates thelength ofthethin filaments; tropomyosin, which stabilizesthin filaments and, inassociationwithtroponin, regulatesbinding ofcalcium ions;M lineproteins(myomesin, M-protein, obscurin),which hold thickfilaments in registeratthe M line; myosin-binding protein C,which contributes to normal assemblyofthick filaments and interacts with titan; and two proteins (desmin and dystrophin) that anchor sarcomeres into the plasma membrane.

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